Beyond Oncogenes: How Ras–MAPK Variants Influence Neurodevelopment

P. Havlickova, A. Koutska, I. Kuta Smatanova, M. Fenckova

Faculty of Science, University of South Bohemia in Ceske Budejovice, Branisovska 1760, Ceske Budejovice, 37005, Czech Republic

fenckm00@prf.jcu.cz

The Ras–MAPK pathway, best known for its role in oncogenesis, is also critical for brain development. Germline mutations in pathway components cause RASopathies—the most common monogenic cause of intellectual disability (ID) and autism spectrum disorder (ASD)—and large-scale sequencing studies continue to uncover numerous additional variants in individuals with neurodevelopmental disorders (NDDs). Most are missense substitutions with uncharacterized functional effects, yet they often cluster within structural “hotspots” that control nucleotide binding, GTP hydrolysis, dimerization, or autoinhibitory regulation [1-3].

Here, large-scale genomic datasets are combined with high-resolution structures of Ras, B-Raf, MEK, and ERK to map these variants and infer their mechanistic consequences. Structural patterns reveal recurrent modes of dysregulation and show parallels with oncogenic mutations, indicating potential opportunities for targeted drug repurposing.

This structural–functional perspective refines pathogenicity assessment, explains phenotypic variability, and identifies priorities for functional validation. By shifting from purely statistical classification to mechanism-based interpretation, it establishes a framework for developing mutation-specific therapeutic strategies in Ras–MAPK–related neurodevelopmental disease.

1.         Rauen K. A. (2013). The RASopathies. Annual review of genomics and human genetics, 14, 355–369. https://doi.org/10.1146/annurev-genom-091212-153523

2.         Geoffray, M. M., Falissard, B., Green, J., Kerr, B., Evans, D. G., Huson, S., Burkitt-Wright, E., & Garg, S. (2021). Autism Spectrum Disorder Symptom Profile Across the RASopathies. Frontiers in psychiatry, 11, 585700. https://doi.org/10.3389/fpsyt.2020.585700

3.         Wang, T., Kim, C. N., Bakken, T. E., Gillentine, M. A., Henning, B., Mao, Y., Gilissen, C., SPARK Consortium, Nowakowski, T. J., & Eichler, E. E. (2022). Integrated gene analyses of de novo variants from 46,612 trios with autism and developmental disorders. Proceedings of the National Academy of Sciences of the United States of America, 119(46), e2203491119. https://doi.org/10.1073/pnas.2203491119

This work is supported by by a grant from the Czech Science Foundation (grant no. 23-07810S) and an EMBO Installation grant (grant no. IG-5310-2023) to M. Fenckova.